Message Not Delivered: Content loss during delivery
The message could not be delivered because it included content or attachments that needed to be converted and information was lost in the process.
Fixes
contact the recipient(s) to ensure that no undesirable consequences result from information being lost from this message before it was delivered.
If the message had attachments, try sending it again without them, and then try sending each attachment in a separate email. If this error occurs when there are no attachments, check the text of the message for unusual content or characters. If it occurs with a specific type of attachment, consider sending that information in another format.
5.6.4: Official Definition
[RFC3463] (Standards Track) from the The Internet Engineering Task Force (IETF) definition for codes that start with 5.X.X:
A permanent failure is one which is not likely to be resolved by resending the message in the current form. Some change to the message or the destination must be made for successful delivery.
[RFC3463] (Standards Track) from the The Internet Engineering Task Force (IETF) definition for codes that end with X.6.4:
This is a warning sent to the sender when message delivery was successfully but when the delivery required a conversion in which some data was lost. This may also be a permanent error if the sender has indicated that conversion with loss is prohibited for the message.